Examines how microbial and chemical processes interact on the Earth's surface today and have shaped the planet throughout its history. Emphasis will be placed on how the life styles and chemical ingenuity of microorganisms drive key biogeochemical processes including weathering and transformations of carbon, oxygen, sulfur, iron and nitrogen. Towards this goal, major geologic and evolutionary events will be examined through the lens of microbial diversity, metabolic energetics, microbe-mineral interactions, and molecular biomarkers. Recommended prerequisites: GEOL 1180 or MCDB 1150 or GEOL 3320 or EBIO 3400 or ENVS 4160 or EVEN 4484. Same as GEOL 5185, ENVS 4185, and MCDB 4185.