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Publications in VIVO

Klymkowsky, Mike Professor


Research Areas research areas


research overview

  • We work in two areas. The first is traditional cell and molecular biology, focussed on the molecular and cellular behavior of complex macromolecular systems, studied in the context of the developing Xenopus embryo. We are currently exploring the use of ultra-high resolution expansion microscopy to examine processes ranging from muscle formation, ciliogenesis, desmosome assembly and disassembly, and synapse formation. My other area of effort is in how to design, assess, and effectively teach students to understand, appreciate, and make use of chemistry (CLUE) and core biology (biofundamentals). These works, carried out in collaboration with Melanie Cooper at Michigan State, have led to multiple free (open educational resources or OERs) and demonstrated and dramatic improvements in student learning and success. As part of this project we employ the beSocratic web-based formative assessment system to evaluated and encourage student learning.


  • cell and molecular biology of early embryonic development, mesoderm formation, neural crest induction; organ formation; course and curricular design, the evaluation and improvement of biology education


selected publications


courses taught

  • MCDB 4100 Special Topics (Spring 2019)
  • MCDB 4650 Developmental Biology (Spring 2019)
  • MCDB 4840 Upper-Division Independent Study (Spring 2019)
  • MCDB 7840 Graduate Independent Study (Spring 2019)
  • FYSM 1000 First Year Seminar (Fall 2018)
  • MCDB 4990 Honors Thesis (Fall 2018)
  • MCDB 7840 Graduate Independent Study (Fall 2018)
  • MCDB 2222 Core Concepts in Biology II: Genes, Genetics and Phenotypes (Spring 2018)
  • MCDB 4100 Special Topics (Spring 2018)
  • MCDB 4840 Upper-Division Independent Study (Spring 2018)
  • MCDB 7840 Graduate Independent Study (Spring 2018)


awards and honors

International Activities

global connections related to teaching and scholarly work (in recent years)

Other Profiles